Elements of courtship song have been implicated as means by which females select conspecific mates in several Drosophila species (eg, Ewing and Bennet-Clark, 1968; Cowling and Burnet, 1981; Kyriacou and Hall, 1982; Hoikkala and Lumme, 1987; Crossley and Bennet-Clark, 1993; Tomaru et al., 1995). Courtship songs differ greatly among closely related Drosophila species (eg, Chang and Miller, 1978; Cowling and Burnet, 1981; Geng et al., 1989; Hoikkala et al., 1994; Ritchie and Gleason, 1995), suggesting that their rate of