We sequenced multiple genes from the enigmatic genera Bhagadatta Moore, 1898, Kumothales Overlaet, 1940 and Harmilla Aurivillius, 1892 (Nymphalidae, Limenitidinae) and analysed them together with a large published dataset. We find that Bhagadatta is sister to the genera Cymothoe Hübner, 1819+ Harma Doubleday, 1848, and that Kumothales is sister to these three. Harmilla is nested within the genus Euriphene Boisduval, 1847. We thus transfer Kumothales and Bhagadatta to the tribe Cymothoini , and we synonymise Harmilla syn. nov. with Euriphene .